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AP Biology Notes

8.3.5 Exponential growth and rmax

AP Syllabus focus:

‘When reproduction is unconstrained, populations can grow exponentially at a maximum per capita growth rate, rmax.’

Exponential population growth describes how a population increases when conditions are ideal. In AP Biology, focus on the meaning of unconstrained reproduction, the shape of exponential (J-shaped) growth, and the biological interpretation of rmaxr_{\max}.

What exponential growth means in population ecology

Core idea: unconstrained reproduction

Exponential growth occurs when a population experiences few or no limiting factors, so individuals reproduce at their physiological potential and survival is high.

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Human population size plotted against time shows the hallmark accelerating pattern associated with exponential growth: the curve becomes progressively steeper as NN increases. The multiple regional lines also reinforce that the same general growth form can occur in different populations under broadly favorable conditions. Source

Exponential growth: Population growth in which the rate of increase is proportional to the current population size, producing a J-shaped curve when population size is plotted against time.

Because the growth rate scales with population size, larger populations add more individuals per unit time than smaller populations, even if each individual’s reproductive output stays the same.

Key conditions that allow exponential growth

Exponential growth is most plausible over short time spans or in newly available habitats. It is associated with:

  • Abundant resources (food, water, nutrients, space)

  • Low competition (few individuals using the same resources)

  • Low predation/parasitism/disease

  • Favourable abiotic conditions (temperature, moisture, pH within tolerance)

  • High reproductive output and/or short generation time in the organism

The parameter rmaxr_{\max} and per capita growth rate

Interpreting rmaxr_{\max}

In exponential growth models, the relevant rate is the maximum possible per-individual contribution to population growth under ideal conditions.

rmaxr_{\max} (maximum per capita growth rate): The highest possible rate at which a population can increase per individual per unit time when reproduction and survival are not limited by environmental resistance.

Biologically, rmaxr_{\max} reflects life-history traits such as age at first reproduction, number of offspring produced, frequency of reproduction, and survivorship patterns.

Per capita vs total population growth

Population increase can be described at two levels:

  • Per capita growth rate: the average contribution of each individual to population change per unit time

  • Total growth rate: the actual increase in number of individuals per unit time for the whole population

In exponential growth, even if the per capita rate stays constant at (or near) rmaxr_{\max}, the total number added per unit time accelerates as NN increases.

Mathematical model of exponential growth (conceptual and quantitative)

A common AP Biology representation links growth rate directly to population size:

dNdt=rmaxN \frac{dN}{dt} = r_{\max}N

dNdt \frac{dN}{dt} = change in population size per unit time (individuals per time)

rmax r_{\max} = maximum per capita growth rate (per time)

N N = population size (individuals)

This form highlights the key proportionality: if NN doubles, the instantaneous growth rate dNdt\frac{dN}{dt} also doubles, assuming the population remains unconstrained.

Reading exponential growth graphs

When interpreting population size vs time:

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The same exponential growth data are shown using different axis scalings: a linear (arithmetic) y-axis produces a J-shaped curve, while a semi-log plot converts the exponential phase into a straight line. This visualization helps justify why a constant per-capita growth process can look “sudden” later in time—because NN is larger—even though the underlying rate parameter remains constant under unconstrained conditions. Source

  • Exponential growth produces a J-shaped curve

  • Early in growth, increases may look small because NN is small

  • Later, growth appears “sudden” because the same per capita rate acts on a much larger NN

When interpreting per capita growth:

  • Under unconstrained conditions, per capita growth is approximately constant and near rmaxr_{\max}

  • If the per capita growth rate decreases with increasing NN, conditions are no longer unconstrained (and the growth pattern is no longer purely exponential)

Biological significance and limits (within the exponential framework)

Why exponential growth matters

Exponential growth models are used as a baseline expectation for what populations can do under ideal conditions. They are especially relevant to:

  • Colonising populations entering a new habitat

  • Recovering populations after a disturbance that reduced density

  • Microbial populations in resource-rich culture conditions for short periods

What determines the magnitude of rmaxr_{\max}

Different species have very different rmaxr_{\max} values due to:

  • Generation time (shorter generation time tends to raise rmaxr_{\max})

  • Fecundity (more offspring per reproductive event tends to raise rmaxr_{\max})

  • Reproductive frequency (more frequent reproduction tends to raise rmaxr_{\max})

  • Survivorship (higher survival to reproductive age can raise effective growth)

If environmental conditions remain ideal, a population can sustain growth near rmaxr_{\max}; if conditions shift, the realised per capita growth rate falls below rmaxr_{\max}.

FAQ

Plot $\ln(N)$ against time during the early, unconstrained phase.

The gradient approximates the per capita growth rate, which may approach $r_{\max}$ if conditions are truly ideal.

Not always.

$r_{\max}$ refers specifically to the maximum possible per capita growth rate under ideal conditions, whereas $r$ can refer to the realised per capita growth rate in the measured environment.

Demographic randomness can dominate at low $N$.

Chance variation in births/deaths, difficulty finding mates, or skewed sex ratios can reduce realised growth below $r_{\max}$.

For exponential growth, doubling time depends on the per capita growth rate.

Higher per capita growth rates imply shorter doubling times; the relationship is commonly expressed using $\ln(2)$ in exponential-growth mathematics.

No.

$r_{\max}$ is defined as the maximum potential per capita growth rate under ideal conditions, so it is non-negative; negative realised growth reflects non-ideal conditions, not $r_{\max}$.

Practice Questions

State what is meant by exponential population growth and identify the parameter rmaxr_{\max} in the model dNdt=rmaxN\frac{dN}{dt} = r_{\max}N. (2 marks)

  • Exponential growth: growth rate proportional to current population size / J-shaped increase over time (1)

  • rmaxr_{\max}: maximum per capita growth rate under unconstrained/ideal conditions (1)

A population enters a new habitat with abundant resources. Using the model dNdt=rmaxN\frac{dN}{dt} = r_{\max}N, explain how and why the pattern of population increase changes over time during exponential growth, and describe what biological traits influence rmaxr_{\max}. (5 marks)

  • Under unconstrained conditions, per capita growth is (approximately) constant at/near rmaxr_{\max} (1)

  • As NN increases, total increase per unit time rises because growth rate is proportional to NN (1)

  • Population size vs time shows a J-shaped curve/accelerating increase (1)

  • rmaxr_{\max} influenced by generation time/age at first reproduction (1)

  • rmaxr_{\max} influenced by fecundity and/or reproductive frequency and/or survivorship to reproduction (1)

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